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Theodoxus fluviatilis (Linnaeus, 1758)
– second part –
Akidalia Fountain (the Spring of the Three Charites), at the foot of the Acropolis hill, Orchomenós, Bœotia, Central Greece. 9mm.
Aréthousa Spring, south of Áyos Stéfanos chapel on the eastern shore of the eponymous bay, south of Chalkída, western Évvia. 5,3mm.
The back of this dark shell (at right and above - 5,3mm) has less black than on the other sides, and shows an abrupt transition to a dotted pattern. At left and below: another specimen with no black coating (4,5mm).
Another image of the 4,5mm specimen from Aréthousa.
Same spring. 5,25-6,05mm.
On stone, Makrisaíos brook, Livadeiá, Boeotia, Central Greece. 6,35mm.
On stones, Krya Springs, Livadeiá. 5,5mm.
Same spot. 5mm.
Same spot. 5mm.
Same spot. 5,8mm.
Pineiós river, in the Vale of Témpi, Dímos Tembón, west of Larissa, northern Thessaly, E. Greece. 7mm.
Neritina heldreichi von Martens, 1879 from SW. Turkey, Crete.
 
Two morphs are recognized in Turkey: “heldreichi” s.s, large, restricted to Eğirdir and Beyşehir Lakes, in Isparta Province; “heldreichi fluvicola” Schütt & Şeşen 1992, smaller, living in streams and springs in Antalya, Burdur, Afyon, Izmir, Manisa, Aydin, Denizli and Isparta.
 
« This species is generally very close to N. fluviatilis, but with the small size of the spire and the mouth narrowed anteriorly, it strays from it: on top of this comes the striking colouring of the mouth. From T. von Heldreich, the Berlin Museum received a Neritine, similar in shape, but much smaller […] with a more jagged design on a more yellow-brown background, also provided with three darker zones, coming from the salt source Almyros at Herakleion in Candia (Crete). » – E. von Martens: “Die Gattung Neritina”, Systematisches Conchylien-Cabinet Bd.2:Abt.10, Nürnberg 1879, p.225.
 
Above: specimens collected on rocks and stones in Manavgat river, Hisarönü Parkı, near the ruins of the castle, Manavgat, Antalya province, S. Turkey. 5,5-7,5mm. These shells bear the jagged pattern seen on the specimens given by von Helreich. This pattern is very close to that of the fluviatilis pictured on top of the previous page. Indeed, heldreichi is now recognized as a synonym of fluviatilis.
Two small specimens from Beyşehir Lake. 4,5-5,35mm.
Same spot. 6,5mm.
Another specimen from Hisarönü Parkı. 6,2mm.
The “Almyros” salt source von Martens is talking about is a pool located west of Heraklion; this is a brackish karst spring that is at the origin of a small coastal stream that flows into the sea on the territory of Skafidáras, Gázi. This spring is one of the major water supplies of the northern Crete.

A “dniestroviensis”, or “heldreichi” subvariant “fluvicola”, a little “intextus” and somewhat “marmoreus”, from the Almyros of Gázi. 3mm. Dogs and cats may be less variable than Theodoxus fluviatilis.
Specimens on stones, at the brackish water spring of Gázi. Notice the « three darker zones » described by von Martens on the specimens collected by Theodor von Heldreich. The latter was a german botanist, specialist of the greek flora.
 
The freshwater of Gazí emergence comes mainly from the Psilotiris Mountains, and reaches Almyros in about nine hours only. It is stopped by the Tylissos-Krousónas fault, which separates the uplifted Psilotiris system from the collapsed Heraklion plain, and receives here some sea water. « At this time, the most satisfying assumption is the digging of an important aquifer, deep below the current sea-level, during the regression of the end of the Miocene. » – R. Maire: La Haute Montagne calcaire, Presses Universitaires de Bordeaux 1990, chapter III “Les hauts karsts de Crète”, section 155.
Corroded adult collected at the outfall. 6,3mm.
Same spot. 5,8-6mm.
Striped specimens from the same spot. 5-5,8mm.
Another Heldreich-style patterned shell from the emergence: compressed jagged stripes with three dark zones. 5,8mm.
Many variations are present also at the Almyros spring of Georgioúpoli, east of Chaniá, NW. Crete. Original picture provided by edpl for iNaturalist – (CC BY-NC).
The Almyros flood of Georgioúpoli, immediately downstream the emergence. This river seems to be corrosive enough to endanger the population of fluviatilis that lives here. Old specimens do not exist, and the apical area is often very destroyed in the adults. Examples below:
Variations of shape in young adults from the emergence. Notice how the older parts of the shells are attacked. This phenomenon, often encountered in all the species of Nerites, is very pronounced in the shells found at the emergences of Georgioúpoli and Gázi. Sizes ≈ 4,2mm.
Colour set found at the spring. 3,6-4mm.
On stones in the wetland.
Growth set from the wetland downstream the spring. The chemical erosion of their shells does not allow these snails to reach old age. Hence the small sizes: 1,3-1,5-3-3,3mm.
Specimens from the lower part of the Almyros, collected in marine sediments near the river mouth. 2,6-3,4mm.
Same spot. 2,5-4,6mm. These shells bear a pattern close to the variant “dniestroviensis” Put’, 1957.
Kalyvaki river, in the northwest of Almyros harbour. On stone on the left bank at 50m upstream the outlet.
Kalyvaki. The genus Theodoxus appreciates the organic film that is found on aquatic plants, but the depth at which the latter live here, in an almost pure marine water, forbids the snails to reach them; this is the reason why they focus on the stones and pebbles that lay on shallower bottoms, in freshwater.
Kalyvaki river, on the rocky western bank. 5,6mm.
Yellow periostracum.
Pattern variations in Kalyvaki. 5-5,5mm.
In this river, the individuals bear a conspicuous extension in the continuation of the rib-shield.
Same spot. 4,3-4,9mm.
Same spot. 5-5,2mm.
Despite recurring summer water stress, the lake Kournas, in the south of Georgioúpoli, hosts another population. Here, a subadult from the outflow in may 2023. 4,95mm.

— return to part 1 or go to part 3, or back to Neritininae —